The prothorax is the foremost of the three segments in the
thorax of an
insect, and bears the first pair of legs. Its principal
sclerites (exoskeletal plates) are the pronotum (
dorsal), the prosternum (
ventral), and the propleuron (
lateral) on each side. The prothorax never bears wings in extant insects, though some fossil groups possessed wing-like projections. All adult insects possess legs on the prothorax, though in a few groups (e.g., the
butterfly family
Nymphalidae) the forelegs are greatly reduced. In many groups of insects, the pronotum is reduced in size, but in a few it is hypertrophied, such as in all
beetles (
Coleoptera), in which the pronotum is expanded to form the entire dorsal surface of the thorax, and most
treehoppers (family
Membracidae, order
Hemiptera), in which the pronotum is expanded into often fantastic shapes that enhance their
camouflage or
mimicry.
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